Nothing reacts instantly, but let's assume
the treadmill matches speed almost instantly, like in a millisecond (1/1000 of a second) and the plane moved forward at 5mph:
1st millisecond plane 5mph -treadmill matches.
2nd millisecond plane's tire is forced to 10mph -treadmill matches.
3rd millisecond plane's tire is forced to 15mph -treadmill matches.
4th millisecond plane's tire is forced to 20mph -treadmill matches.
After a full second the wheels are spinning at 5,000mph.
Two forces that are plied against each other, unconstrained, compounding each other instantly, would become incredibly volatile within seconds.